diff --git a/oauth2_http/java/com/google/auth/oauth2/ComputeEngineCredentials.java b/oauth2_http/java/com/google/auth/oauth2/ComputeEngineCredentials.java
index fd7260534..2e0c321d1 100644
--- a/oauth2_http/java/com/google/auth/oauth2/ComputeEngineCredentials.java
+++ b/oauth2_http/java/com/google/auth/oauth2/ComputeEngineCredentials.java
@@ -657,6 +657,12 @@ public Builder setUniverseDomain(String universeDomain) {
       return this;
     }
 
+    @CanIgnoreReturnValue
+    public Builder setQuotaProjectId(String quotaProjectId) {
+      super.quotaProjectId = quotaProjectId;
+      return this;
+    }
+
     public HttpTransportFactory getHttpTransportFactory() {
       return transportFactory;
     }
diff --git a/oauth2_http/java/com/google/auth/oauth2/GoogleCredentials.java b/oauth2_http/java/com/google/auth/oauth2/GoogleCredentials.java
index a49cf9b2f..54d708103 100644
--- a/oauth2_http/java/com/google/auth/oauth2/GoogleCredentials.java
+++ b/oauth2_http/java/com/google/auth/oauth2/GoogleCredentials.java
@@ -481,7 +481,9 @@ protected Builder() {}
     protected Builder(GoogleCredentials credentials) {
       super(credentials);
       this.quotaProjectId = credentials.quotaProjectId;
-      this.universeDomain = credentials.universeDomain;
+      if (credentials.isExplicitUniverseDomain) {
+        this.universeDomain = credentials.universeDomain;
+      }
     }
 
     protected Builder(GoogleCredentials.Builder builder) {
diff --git a/oauth2_http/javatests/com/google/auth/oauth2/ComputeEngineCredentialsTest.java b/oauth2_http/javatests/com/google/auth/oauth2/ComputeEngineCredentialsTest.java
index 80268bb1e..dc307d518 100644
--- a/oauth2_http/javatests/com/google/auth/oauth2/ComputeEngineCredentialsTest.java
+++ b/oauth2_http/javatests/com/google/auth/oauth2/ComputeEngineCredentialsTest.java
@@ -60,6 +60,7 @@
 import java.util.Arrays;
 import java.util.Collection;
 import java.util.Collections;
+import java.util.Date;
 import java.util.List;
 import java.util.Map;
 import java.util.Queue;
@@ -361,6 +362,19 @@ public void hashCode_equals() throws IOException {
     assertEquals(credentials.hashCode(), otherCredentials.hashCode());
   }
 
+  @Test
+  public void toBuilder() {
+    ComputeEngineCredentials credentials =
+        ComputeEngineCredentials.newBuilder()
+            .setHttpTransportFactory(new MockMetadataServerTransportFactory())
+            .setQuotaProjectId("quota-project")
+            .build();
+
+    ComputeEngineCredentials secondCredentials = credentials.toBuilder().build();
+
+    assertEquals(credentials, secondCredentials);
+  }
+
   @Test
   public void serialize() throws IOException, ClassNotFoundException {
     MockMetadataServerTransportFactory serverTransportFactory =